Update, 4:26 p.m. eastern: Deadline has amended their earlier Friday post, and are now saying that John C. Reilly's Guardians of the Galaxy role is indeed Rhomann Dey, a leader of the Nova Corps — and not Ronan the Accuser, as they previously reported.
This is the same role that Hitfix reported Reilly was offered back in late May. At the time, the site described the part as essentially the Agent Coulson of the Guardians films — "He's the human agent who serves as a go-between, keeping the Guardians in line and reporting to S.H.I.E.L.D. about their activities."
Original story: A new Guardians of the Galaxy role has been confirmed by Deadline, with the site reporting that actor John C. Reilly is playing long-running Marvel character Ronan the Accuser.
This combines two previously unconfirmed tidbits surrounding the movie; word from late last month that Ronan would be a major villain in the film, and that Reilly would be in the movie as Nova Corps leader Rhomann Dey. Curiously, Deadline lists Reilly's role as "Ronan 'The Accuser' Dey," a last name not previously associated with the character in the comics.
Ronan, introduced by Stan Lee and Jack Kirby in 1967's Fantastic Four #65, is a prominent member of the Marvel Universe's Kree alien race, and while originally depicted as a villain, he's recently been portrayed as a hero in stories like Annihilators.
Reilly is known for both comedic and dramatic roles, ranging from Boogie Nights to Step Brothers, to voicing the title character in Disney's Wreck-It Ralph. He was nominated for an Academy Award for his role in 2002's Chicago.
Several more actors have been reported to be part of Guardians of the Galaxy with no characters officially attached, including Benicio del Toro, Glenn Close, Karen Gillan and Lee Pace.
Guardians of the Galaxy, written and directed by James Gunn, is scheduled for release on Aug. 1, 2014.